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Data associated with selected splice site

Sequence

 atgc   intronic sequence     ATGC   exonic sequence

gtgagtcccctacttatttcaacacatttgatgctcatttttctccattatattttaagtatttttttgacaagtgctgtttaattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG

Basic information

species Zea mays
transcript GRMZM2G421857_T04
intron # 17
splice site 3'
intron type U2

Orthologous splice sites


 atgc   intronic sequence     ATGC   exonic sequence


upper sequence: GRMZM2G421857_T02 (Zea mays), 3'ss of exon 16
lower sequence: LOC_Os02g07870.1 (Oryza sativa), 3'ss of exon 16
gtgagtcccctacttatttcaacacatttgatgctcatttttctccattatatttta-agtatttttttgacaagtgctgtttaattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG
||||||     ||  |||| |       |  | || | |   ||   | ||| |  | | ||  |  ||||  | | |||||  || | | |||||||||||||  ||||||||||||||||| |||||||||||||||||||||||||| ||||||||||||||||  |||
gtgagtgttaaacccatttga-------tagttct-actaacctgtgtcataatccatattaactaattgatcaattctgttgcatgaaacagATATGATAAGTATTGCCCATTCTACAAATCTGTTTGGATGATGCGCAACATTATTCATTTCAACACATTGGCAACCCAG

upper sequence: GRMZM2G421857_T02 (Zea mays), 3'ss of exon 16
lower sequence: GLYMA07G00430.1 (Glycine max), 3'ss of exon 17
gtgagtcccctacttatt--tcaacacatttgatgctcatttttctccattatattttaagtatttttttgacaagtgctgtttaattat-atagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG
|| ||  ||||  | |||  | |||   ||| |   | ||| |  |  ||| | || |      | | |||  |  | |||| |   | | ||||||||||||| || ||||| |||||||| || ||||||||||||||||||||||| || ||| |||  ||||||||||||
gtaagggcccttgtaattggttaacttctttaacatttattatatt--attgtgttct------tatgttggtaccttctgtgtggctgtgatagATATGATAAATTTTGCCCCTTCTACAAGTCTGTTTGGATGATGCGCAACATTATCCATTTCTACAATTTGGCAAATCAG

upper sequence: GRMZM2G421857_T02 (Zea mays), 3'ss of exon 16
lower sequence: GLYMA08G23990.1 (Glycine max), 3'ss of exon 17
gtgagtcccctacttatt--tcaacacatttgatgctcatttttctccattatattttaagtatttttttgacaagtgct-gtttaattat-atagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG
|| ||   |||  | |||  | |||   ||| ||    ||| |  |    | | | ||   | || | |||  |  |  | || |   | | |||||||||| || |||||||| |||||||| || ||||||||||||||||||||||| || ||| |||  ||||||||||||
gtaagggaccttgtaattggttaacttctttaatatgtattatattattgtttttatttgttcttatgttggtacctttttgtgtggctgtgatagATATGACAAATTCTGCCCCTTCTACAAGTCTGTTTGGATGATGCGCAACATTATCCATTTCTACAATTTGGCAAATCAG

upper sequence: GRMZM2G421857_T02 (Zea mays), 3'ss of exon 16
lower sequence: AT1G78900.2 (Arabidopsis thaliana), 3'ss of exon 17
--gtgagtcccctacttatttcaacaca-tttgatgctcatttttctccattatattttaagtatttttttgacaagtgctgtttaattat---atagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG
    || |      ||   |||       |||  | |||  | ||   |  |  |       |    | ||||    |   | | |||| |   | |||||||| || ||||| || |||||||| ||||| ||||||||||| |||||||| || ||| |||   | || || |||
gtaagatttgttggctcccttcgttttggtttagtactctctctttctctctcaacgggttattcactcttgaaccttttggatgaattttttgacagATATGACAAATTCTGTCCTTTCTACAAGTCCGTGTGGATGATGCGTAACATTATCCATTTCTACAACCTAGCCAACCAG

upper sequence: GRMZM2G421857_T02 (Zea mays), 3'ss of exon 16
lower sequence: Vv09s0054g00890.t01 (Vitis vinifera), 3'ss of exon 17
----------------------------------------gtgagtcccctacttatttcaacacatttgatgctcatttttctccattatattttaagtatttttt-----tgacaagtgc----tgtttaattat-atagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG
                                         |        |  ||| ||         | ||| ||   ||| |    |    |  || |  |||||     | | |  ||       | |   | | | |||||||||||||||||||| |||||||| || ||||||||||||||||| ||||| || ||| | |  ||||| ||||||
gtaagtcccagattaagtttttgattcaaattttcgtcctttttaatttttgtttacttactatttccttatgttcccctttttatgctggtctagaaattctttttccacttaatatatgtatctcatgtggctgtgacagATATGATAAGTTCTGCCCCTTCTACAAGTCGGTTTGGATGATGCGCAATATTATCCATTTCTATAATTTGGCCAATCAG

upper sequence: GRMZM2G421857_T02 (Zea mays), 3'ss of exon 16
lower sequence: PP1S44_102V6.1 (Physcomitrella patens), 3'ss of exon 17
--gtgagtcccctacttatttcaacacatttga--tgctcatttttctccattatatttt---aagtatttttttgacaagtgctgtttaattata-tagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG
   ||          | | |  |       |    ||   |  |  ||  | | |  |||   | ||   | | ||||| | |||  | |    |   || |||||||| || |||||||| || || || || |||||||||||||||||| | |||||||||||  | || ||||||
cgatgcactttgccttgaatagattgggggtagagtgtcaacataccttgaatgtgattttgcatgtgcgtatctgacatgggcttgtcatgggtggcagTTATGATAAATTTTGCCCATTTTATAAGTCGGTGTGGATGATGCGCAACATTGTCCACTTCAACACTCTTGCCAATCAG

upper sequence: GRMZM2G421857_T02 (Zea mays), 3'ss of exon 16
lower sequence: PP1S10_201V6.1 (Physcomitrella patens), 3'ss of exon 17
--gtgagtcccctacttatttcaacacatttgatgctcatt-tttctccattatat-----tttaagtatttttttgacaagtgctgtttaattata-tagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG
   ||  |  | |   || |  |   || |||  | | |    ||  | | | |||     |||   |     | |||   | | |  | |    |   || ||||||||||||||||||||||| ||||| || |||||||||||||| ||| | ||||| |||||  | || ||||||
ttatgcatttcgt-tataatagattgcagttgtggatgacagcttgcctaatctatggatgtttctatccaagtctgatgtgggttcctcatgggtggcagTTATGATAAGTTCTGCCCATTCTATAAATCAGTATGGATGATGCGCAATATTGTCCACTTTAACACTCTTGCCAATCAG

upper sequence: GRMZM2G421857_T02 (Zea mays), 3'ss of exon 16
lower sequence: PP1S8_312V6.1 (Physcomitrella patens), 3'ss of exon 17
--------------------------------gtgagtcccctacttatttcaacacatttgatgctcatttttctccattatattttaagtatttttttgacaagtgctgtttaattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG
                                ||  ||    ||  | |||      |||   ||  | | ||   | ||| | | ||      | | |      || || |   |    | || ||||| || ||||||||||||||||| || || ||||||||||| || ||| | |||||||||||  | |||||||||
gtgagctttgttgaaattttgttgtgtaggatgtttgttgggtagatgtttttcattattctgtgtccctgttcgacgattgtgtgtttctccatgtctgattcggtcctctcatacgtgacagTTATGACAAATTCTGCCCATTCTACAAGTCTGTGTGGATGATGCGTAATATTGTCCACTTCAACACTCTTGCAAATCAG

upper sequence: GRMZM2G421857_T02 (Zea mays), 3'ss of exon 16
lower sequence: EFJ33015 (Selaginella moellendorffii), 3'ss of exon 16
gtgagtcccctacttatttcaacacatttgatgctcatttttctccattatattttaagtatttttttgacaagtgctgtttaatta-tatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG
                                                  | ||    |  |  | |||   |  |||    |  |  |  ||||| |||||||| || || |||||||| ||||||||||||||||| |||||  |||| |||  |    || | | ||||
---------------------------------------------gtacgtgttccgggcttgctcttgtttatcgctaacaatctgctccagATACGATAAGTTTTGTCCGTTCTACAAGTCCGTTTGGATGATGCGAAACATCGTTCATTTCTGCGACCTGTCCACTCAG

upper sequence: GRMZM2G421857_T02 (Zea mays), 3'ss of exon 16
lower sequence: EFJ30330 (Selaginella moellendorffii), 3'ss of exon 17
gtgagtcccctacttatttcaacacatttgatgctcatttttctccattatattttaagtatttttttgacaagtgctgtttaatta-tatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG
                                                  | ||    |  |  | |||   |  |||    |  |  |  ||||| |||||||| || || |||||||| ||||||||||||||||| |||||  | || |||  |    || | | ||||
---------------------------------------------gtacgtgttccgggcttgctcttgtttatcgctaacaatctgctccagATACGATAAGTTTTGTCCGTTCTACAAGTCCGTTTGGATGATGCGAAACATCGTCCATTTCTGCGACCTGTCCACTCAG

Mapped EST sequences

Showing partial alignments of ESTs and genomic sequences. See full alignments


 ATGC     EST sequence
 ATGC     genomic sequence (exon)
 ATGC     genomic sequence (truncated intron)


EST: gi|32850183|gb|CD989864.1|CD989864
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|32921243|gb|CF026055.1|CF026055
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|67026160|gb|CO454909.1|CO454909
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCTGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|32845901|gb|CD985582.1|CD985582
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|32922059|gb|CF026871.1|CF026871
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|24774168|gb|CA409422.1|CA409422
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|8368089|gb|BE051034.1|BE051034
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCTGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|32921648|gb|CF026460.1|CF026460
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|211529162|gb|FL123428.1|FL123428
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|6020503|gb|AW065431.1|AW065431
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|32849884|gb|CD989565.1|CD989565
EST:     ACAGCAAAGCTTNTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|211538183|gb|FL123481.1|FL123481
EST:     ACAGCAAAGCTTTTGAGARAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|32846431|gb|CD986112.1|CD986112
EST:     ACAGCAAAGCTTTTGACAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGAATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|24773770|gb|CA409024.1|CA409024
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|32846734|gb|CD986415.1|CD986415
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|8368090|gb|BE051035.1|BE051035
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCTGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|211156007|gb|FL123542.1|FL123542
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCATCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|8665370|gb|BE186186.1|BE186186
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCTGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|40334940|gb|CK369010.1|CK369010
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|22542087|gb|BU092525.1|BU092525
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|6227177|gb|AW155706.1|AW155706
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|32921742|gb|CF026554.1|CF026554
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|32920116|gb|CF024928.1|CF024928
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|32921546|gb|CF026358.1|CF026358
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
EST: gi|60352649|gb|DN219622.1|DN219622
EST:     ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCC                         ATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT
genomic: ACAGCAAAGCTTTTGAGAGAAGATTATTTGGCACAAAATGCATTCACCCCgtgagtcccc ... aattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACAT


 atgc   intronic sequence     ATGC   exonic sequence

Intronic sequence truncated to 55 bases.

tttttctccattatattttaagtatttttttgacaagtgctgtttaattatatagATATGATAAGTTCTGCCCATTCTACAAATCCGTTTGGATGATGCGCAACATTATTCACTTCAACACATTGGCAAATCAG
         attatattttaagtat  TA-rich tract